See also


Titova AV, Straw AD. Contradictory behavioral effects of neuronal perturbations on behavioral responses to linearly polarized light in freely walking Drosophila. bioRxiv (2024) doi:10.1101/2024.03.15.584848
Vo-Doan TT, Titov VV, Harrap MJM, Lochner S, Straw AD. High Resolution Outdoor Videography of Insects Using Fast Lock-On Tracking. bioRxiv (2023) doi:10.1101/2023.12.20.572558 | Movie 1. High-speed video of bumble bee | Movie S5. Quadcopter-based bee tracking
Vo-Doan TT, Straw AD. Millisecond insect tracking system. arXiv (2020) doi:10.48550/arXiv.2002.12100 | arXiv:2002.12100 | Video on YouTube
Poehlmann A*, Soselisa S*, Fenk LM, Straw AD. A unifying model to predict multiple object orienting behaviors in tethered flies. bioRxiv (2018) doi:10.1101/379651

Peer reviewed journal articles


Wittmann K, Ibrahim MG, Straw AD, Klein A-M, Staab M. Monitoring fast moving animals – building a customized camera system and evaluation toolset. Methods in Ecology and Evolution (2024) doi:10.1111/2041-210X.14322 Data at Code at


Titova AV, Kau BE, Tibor S, Mach J, Vo-Doan TT, Wittlinger M, Straw AD. Displacement experiments provide evidence for path integration in Drosophila. Journal of Experimental Biology (2023) doi:10.1242/jeb.245289 Preprint on bioRxiv.
Straw AD, Pieters RPM, Muijres FT. Real-Time Tracking of Multiple Moving Mosquitoes. Cold Spring Harbor Protocols (2023) doi:10.1101/pdb.prot107927 URL:


Kellner MJ*, Ross JJ*, Schnabl J*, Dekens MPS, Matl M, Heinen R, Grishkovskaya I, Bauer B, Stadlmann J, Menéndez-Arias L, Straw AD, Fritsche-Polanz R, Traugott M, Seitz T, Zoufaly A, Födinger M, Wenisch C, Zuber J, Vienna Covid-19 Detection Initiative (VCDI), Pauli A, Brennecke J. A Rapid, Highly Sensitive and Open-Access SARS-CoV-2 Detection Assay for Laboratory and Home Testing. Frontiers in Molecular Biosciences (2022) doi:10.3389/fmolb.2022.801309 Preprint on bioRxiv. []
Alonso San Alberto D, Rusch C, Zhan Y, Straw AD, Montell C, Riffell J. The olfactory gating of visual preferences to human skin and visible spectra in mosquitoes. Nature Communications 13(555) (2022) doi:10.1038/s41467-022-28195-x Preprint on bioRxiv.
Cribellier A, Straw AD, Spitzen J, Pieters RPM, van Leeuwen JL, Muijres FT. Diurnal and nocturnal mosquitoes escape looming threats using distinct flight strategies. Current Biology (2022) doi:10.1016/j.cub.2022.01.036


Straw AD. Review of methods for animal videography using camera systems that automatically move to follow the animal. Integrative and Comparative Biology (2021) doi:10.1093/icb/icab126
Fernández A, Straw A, Distel M, Leitgeb R, Baltuska A, Verhoef AJ. Dynamic real-time subtraction of stray-light and background for multiphoton imaging. Biomedical Optics Express 12(1), 288-302 (2021) doi:10.1364/BOE.403255


Linneweber GA, Andriatsilavo M, Dutta SB, Bengochea M, Hellbruegge L, Liu G, Ejsmont RK, Straw AD, Wernet M, Hiesinger PR, Hassan B. A neurodevelopmental origin of behavioral individuality in the Drosophila visual system. Science 367(6482), 1112-1119 (2020) doi:10.1126/science.aaw7182 []


Dakin R*, Segre PS*, Straw AD, Altshuler DL. Morphology, muscle capacity, skill, and maneuvering ability in hummingbirds. Science 359(6376), 653-657 (2018) doi:10.1126/science.aao7104


Stowers JR*, Hofbauer M*, Bastien R, Griessner J⁑, Higgins P⁑, Farooqui S⁑, Fischer RM, Nowikovsky K, Haubensak W, Couzin ID, Tessmar-Raible K✎, Straw AD✎. Virtual Reality for Freely Moving Animals. Nature Methods 14, 995–1002 (2017) doi:10.1038/nmeth.4399 [FreemoVR website]


Segre PS*, Dakin R*, Read TJG, Straw AD, Altshuler DL. Mechanical constraints on flight at high elevation decrease maneuvering performance of hummingbirds. Current Biology 26(24), 3368-3374 (2016) doi:10.1016/j.cub.2016.10.028
Panser K*, Tirian L*, Schulze F*, Villalba S, Jefferis GSXE, Bühler K, Straw AD. Automatic segmentation of Drosophila neural compartments using GAL4 expression data reveals novel visual pathways. Current Biology 26(15), 1943-1954 (2016) doi:10.1016/j.cub.2016.05.052 See also the braincode website. Open-Access Link. Preprint on bioRxiv.


Segre P, Dakin R, Zordan VB, Dickinson MH, Straw AD, Altshuler DL. Burst muscle performance predicts the speed, acceleration, and turning performance of Anna's hummingbirds. eLife 4, e11159 (2015) doi:10.7554/eLife.11159


Fenk LM*, Poehlmann A*, Straw AD. Asymmetric processing of visual motion for simultaneous figure and background responses. Current Biology 24(24), 2913-2919 (2014) doi:10.1016/j.cub.2014.10.042
Bath DE*, Stowers JR*, Hörmann D, Poehlmann A, Dickson BJ✎, Straw AD✎. FlyMAD: Rapid thermogenetic control of neuronal activity in freely-walking Drosophila. Nature Methods 11(7), 756-762 (2014) doi:10.1038/nmeth.2973
Stowers JR, Fuhrmann A, Hofbauer M, Streinzer M, Schmid A, Dickinson MH, Straw AD. Reverse engineering animal vision with virtual reality and genetics. Computer 47(7), 38-45 (2014) doi:10.1109/MC.2014.190
Dell AI, Bender JA, Branson K, Couzin ID, de Polavieja GG, Noldus LPJJ, Perez-Escudero A, Perona P, Straw AD, Wikelski M, Brose U. The role of automated tracking in ecology. Trends in Ecology and Evolution 29(7), 417-428 (2014) doi:10.1016/j.tree.2014.05.004
Fuller SB, Straw AD, Peek MY, Murray RM, Dickinson MH. Flying Drosophila stabilize their vision-based velocity controller by sensing wind with their antennae. PNAS (2014) doi:10.1073/pnas.1323529111


Censi A*, Straw AD*, Sayaman RW, Murray RM, Dickinson MH. Discriminating external and internal causes for heading changes in freely flying Drosophila. PLOS Computational Biology 9(2), 1-14 (2013) doi:10.1371/journal.pcbi.1002891
Zantke J, Ishikawa-Fujiwara T, Arboleda E, Lohs C, Shipany K, Hallay N, Straw AD, Todo T, Tessmar-Raible K. Circadian and circalunar clock interactions in a marine annelid. Cell Reports (2013) doi:10.1016/j.celrep.2013.08.031


Straw AD, Branson K, Neumann TR, Dickinson MH. Multicamera Realtime 3D Tracking of Multiple Flying Animals. Journal of The Royal Society Interface 8(11), 395-409 (2011) doi:10.1098/rsif.2010.0230
Mamiya A, Straw AD, Tómasson E, Dickinson MH. Active and Passive Antennal Movements during Visually Guided Steering in Flying Drosophila. Journal of Neuroscience (2011) doi:10.​1523/​JNEUROSCI.​0498-11.​2011


Straw AD, Lee S, Dickinson MH. The visual control of altitude in flying Drosophila. Current Biology 20(17), 1550-1556 (2010) doi:10.1016/j.cub.2010.07.025
Robie AA, Straw AD, Dickinson MH. Object preference by walking fruit flies, Drosophila melanogaster, is mediated by vision and graviperception. Journal of Experimental Biology (2010) doi:10.1242/jeb.041749
Maimon G, Straw AD, Dickinson MH. Active flight increases the gain of visual motion processing in Drosophila. Nature Neuroscience 13(3), 393-399 (2010) doi:10.1038/nn.2492


Straw AD, Dickinson MH. Motmot, an open-source toolkit for realtime video acquisition and analysis. Source Code Biol Med 4(5) (2009) doi:10.1186/1751-0473-4-5
Fry SN, Rohrseitz N, Straw AD, Dickinson MH. Visual flight speed control in Drosophila melanogaster. J Exp Biol (2009) doi:10.1242/jeb.020768


Straw AD, Rainsford T, O'Carroll DC. Contrast sensitivity of insect motion detectors to natural images. Journal of Vision (2008) doi:10.1167/8.3.32
Straw AD. Vision Egg: An Open-Source Library for Realtime Visual Stimulus Generation. Front Neuroinformatics 2(4) (2008) doi:10.3389/neuro.11.004.2008
Maimon G, Straw AD, Dickinson MH. A simple vision-based algorithm for decision making in flying Drosophila. Current Biology (2008) doi:10.1016/j.cub.2008.02.054
Dickson WB, Straw AD, Dickinson MH. Integrative model of Drosophila flight. AIAA Journal (2008) doi:10.2514/1.29862
Fry SN, Rohrseitz N, Straw AD, Dickinson MH. TrackFly: Virtual reality for a behavioral system analysis in free-flying fruit flies. J Neurosci Meth (2008) doi:10.1016/j.jneumeth.2008.02.016


Straw AD, Warrant EJ, O'Carroll DC. A bright zone in male hoverfly (Eristalis tenax) eyes and associated faster motion detection and increased contrast sensitivity. J Exp Biol (2006) doi:10.1242/jeb.02517


Shoemaker P, O'Carroll DC, Straw AD. Velocity constancy and models for wide-field motion detection in insects. Biological Cybernetics (2005) doi:10.1007/s00422-005-0007-y


Fry SN, Mueller P, Baumann HJ, Straw AD, Bichsel M, Robert D. Context-dependent stimulus presentation to freely moving animals in 3D. J Neurosci Meth (2004) doi:10.1016/j.jneumeth.2003.12.012


* equal contribution

⁑ equal contribution

✎ co-corresponding author

Peer reviewed conference papers


Han S, Censi A, Straw AD, Murray RM. A Bio-Plausible Design for Visual Pose Stabilization. Proceedings of the 2010 IEEE/RSJ International Conference on Intelligent Robots and Systems (IROS), 5679–5686 (2010) doi:10.1109/IROS.2010.5652857


Han S, Straw AD, Dickinson MH, Murray RM. A real-time helicopter testbed for insect-inspired visual flight control. Robotics and Automation, 2009. ICRA '09. IEEE International Conference on, 3055-3060 (2009) doi:10.1109/ROBOT.2009.5152667


Epstein M, Waydo S, Fuller SB, Dickson W, Straw AD, Dickinson MH, Murray RM. Biologically Inspired Feedback Design for Drosophila Flight. American Control Conference, 2007. ACC '07, 3395-3401 (2007) doi:10.1109/ACC.2007.4282971


Dickson W, Straw AD, Poelma C, Dickinson MH. An Integrative Model of Insect Flight Control. 44th AIAA Aerospace Sciences Meeting and Exhibit (2006) doi:10.2514/6.2006-34


Rajesh S, Straw AD, O'Carroll DC, Abbott D. Effect of spatial sampling on pattern noise in insect-based motion detection. Proc. SPIE 5649, Smart Structures, Devices, and Systems II, 811 (March 09, 2005) (2005) doi:10.1117/12.598178
Rajesh S, Straw AD, O'Carroll DC, Abbott D. Effects of compressive nonlinearity on insect-based motion detection. Proc. SPIE 5649, Smart Structures, Devices, and Systems II, 798 (March 09, 2005) (2005) doi:10.1117/12.598177


Shoemaker PA, O'Carroll DC, Straw AD. Implementation of visual motion detection with contrast adaptation. Proc. SPIE 4591, Electronics and Structures for MEMS II, 316 (November 21, 2001) (2001) doi:10.1117/12.449162


Hellekes K, Villalba S, Stowers JR, Graf A, Panser K, Campione E, Straw AD. A dataset of 3D fly (Drosophila melanogaster) flight trajectories to study the role of neuropeptide degradation in visuo-motor behaviors. doi:10.5281/zenodo.29193 link

Online resources

Braincode website - Automatic segmentations of brain regions based on enhancer clustering. link
FlyMAD - the Fly Mind Altering Device - website. link
Models of Visual Fly Motion Detection and Behavior. link

Software and Hardware

Strand Camera - low-latency single camera acquisition and tracking software. link
Braid - multi-camera acquisition and tracking software. link
Miriam - open-source isothermal reaction instrument featuring realtime fluorescence readout in a 96-well plate format. link - Automatic image enhancement for Hydroxy naphthol blue (HNB) dyes. link
FreemoVR - Virtual Reality for Freely Moving Animals. link
Flydra - multi-camera acquisition and tracking software. link
BUI Backend - library to write user interfaces in the browser. link
VisonEgg - simple visual stimulus generation library (Python). link
PyMVG - Python Multiple View Geometry (Python). link
Models of Visual Fly Motion Detection and Behavior (Python). link
Neuron catalog web application (Javascript/Coffeescript). link
FlyMAD - the Fly Mind Altering Device source code (Python). link
Motmot Camera Utilities (Python/C/C++). link